Rhizostomeae

Order Rhizostomeae Cuvier 1799

Characteristics of the Order (from Kramp 1961)

Scyphomedusae with umbrella margin cleft into lappets; without marginal tentacles; without a central mouth opening, but with numerous mouths upon eight adradial, fleshy, branched, arm-like appendages arising from the centre of the subumbrella; with rhopalia between marginal clefts.

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Based largely on Kramp (1961), Mianzan & Cornelius (1999)

Collections: M = morphology, D = DNA.

Type = *

Family (8)	Genus (26)	Species (94) [91 morphological + 3 molecular]	Collections
Cassiopeidae L. Agassiz 1862	Cassiopea Péron & Lesueur 1810	andromeda(Forskål 1775) *	Bermuda (D), Hawaii (D, M), Florida (D), Red Sea (D)
		depressa Haeckel 1880
		frondosa (Pallas 1774)	Florida (D), Panama (D)
		medusa Light 1914
		mertensi Brandt 1838
		ndrosia Agassiz & Mayer 1899
		ornata Haeckel 1880	Fiji (D), Kakaban (D, M), Palau (D, M)
		picta Vanhöffen 1888
		vanderhorsti Stiasny 1922
		sp. 1 Holland et al. (2004)	Queensland (D)
		sp. 2 Holland et al. (2004)	New Guinea (D)
		sp. 3 Holland et al. (2004)	Hawaii (D, M), New Guinea (D)
Catostylidae Gegenbaur 1857	Acromitoides Stiasny 1921	purpurus (Mayer 1910)
		stiphropterus (Schultze 1897)
	Acromitus Light 1914	flagellatus (Maas 1903)	Taiwan (D)
		hardenbergi Stiasny 1934
		maculosus Light 1914 *
		rabanchatu Annandale 1915
		tankahkeei Light 1924
	Catostylus L. Agassiz 1862	cruciatus (Lesson 1830)
		mosaicus (Quoy & Gaimard 1824) *		Queensland (D), New South Wales (D, M), Tasmania (D, M), Victoria (D, M)
		ornatellus (Vanhöffen 1888)
		ouwensi Moestafa & McConnaughey 1966
		perezi Ranson 1945
		tagi (Haeckel 1869)
		townsendi Mayer 1915	Taiwan (D)
		tripterus (Haeckel 1880)
		turgescens (Schultze 1911)
		viridescens (Chun 1896)
	Crambione Maas 1903	bartschi (Mayer 1910)
		cooki Mayer 1910
		mastigophora Maas 1903 *
	Crambionella Stiasny 1921	annandalei Rao 1931
		orsini (Vanhöffen 1888) *	Persian Gulf (D)
		stuhlmanni (Chun 1896)
	Leptobrachia Brandt 1838	leptopus (Chamisso & Eysenhardt 1821) *
Cepheidae L. Agassiz 1862	Cephea Péron & Lesueur 1810	cephea (Forskål 1775)	Palau (D), Kwajalein (D)
		coerulea Vanhöffen 1902
		conifera Haeckel 1880
		octostyla (Forskål 1775) *
		sp. Mayer 1915
		sp. Stiasny 1937
		sp. Kramp 1958
	Cotylorhiza L. Agassiz 1862	ambulacrata Haeckel 1880
		erythraea Stiasny 1920
		pacifica Mayer 1915
		tuberculata (Macri 1778) *
	Netrostoma L.S. Schultze 1898	coerulescens Maas 1903
		dumokuroa (Agassiz & Mayer 1899)
		setouchianum (Kishinouye 1902)
		typhlodendrium L.S. Schultze 1898 *
	Polyrhiza L. Agassiz 1862	vesiculosa (Ehrenberg 1835) *
Lychnorhizidae Haeckel 1880	Anomalorhiza Light 1921	shawi Light 1921 *	Philippines (D, M)
	Lychnorhiza Haeckel 1880	arubae Stiasny 1920
		lucerna Haeckel 1880*
		malayensis Stiasny 1920
	Pseudorhiza von Lendenfeld 1882	aurosa von Lendenfeld 1882 *
		haeckeli Haacke 1884
Lobonematidae Stiasny 1921	Lobonema Mayer 1910	mayeri Light 1914
		smithi Mayer 1910 *
	Lobonemoides Light 1914	gracilis Light 1914 *
		robustus Stiasny 1920
		sewelli Rao 1931
Mastigiidae Stiasny 1921	Mastigias L. Agassiz 1862	albipunctatus Stiasny 1920
		andersoni Stiasny 1926
		gracilis (Vanhöffen 1888)
		ocellatus (Modeer 1791)
		pantherinus Haeckel 1880
		''papua'' (Lesson 1830)*	Berau (D, M), Japan (D, M), Palau (D, M),
		roseus (Reynaud 1830)
		siderea Chun 1896
		sp. Rao 1931
	Mastigietta Stiasny 1921	palmipes (Haeckel 1880) *
	Phyllorhiza L. Agassiz 1862	luzoni Mayer 1915
		pacifica (Light 1921)
		peronlesueuri Goy 1990
		punctata von Lendenfeld 1884 *
	Thysanostoma L. Agassiz 1862	flagellatum (Haeckel 1880)
		loriferum (Ehrenberg 1835)
		thysanura Haeckel 1880 *	Palau (D)
	Versuriga Kramp 1961	anadyomene (Maas 1903) *	Palau (D)
Rhizostomatidae Cuvier 1799	Eupilema Haeckel 1880	inexpectata Pages Gili & Bouillon 1992
		scapulare Haeckel 1880 *
	Nemopilema Kishinouye 1922	''nomurai'' (Kishinouye 1922) *
	Rhizostoma Cuvier 1799	luteum (Quoy & Gaimard 1827)
		octopus (Macri 1778)
		pulmo (Macri 1778)*
	Rhopilema Haeckel 1880	esculentum Kishinouye 1891	China (D, M)
		hispidum (Vanhöffen 1888)
		nomadica Galil, Spanier & Ferguson 1990
		rhopalophorum Haeckel 1880 *
		verrilli (Fewkes 1887)
Stomolophidae Haeckel 1880	Stomolophus L. Agassiz 1862	fritillarius Haeckel 1880
		meleagris L. Agassiz 1862 *

See Zoo Bank ([1]) for more information.

Compiled by M.N Dawson, G. Jarms, A.C. Morandini. Update pending.